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tant points of the southern hemisphere, of species which, though distinct, belong to genera exclusively confined to the south, is a more remarkable case. Some of these species are so distinct that we cannot suppose that there has been time since the commencement of the last glacial period for their migration and subsequent modification to the necessary degree.” Mr. Darwin goes on to account for these facts by the probable existence of a rich antarctic flora in a warm period anterior to the last glacial epoch. There are indeed many reasons for thinking that a southern contiment, rich in living forms, once existed. One such reason is the way in which struthious birds are, or have been, distributed around the antarctic region: as the ostrich in Africa, the rhea in South America, the emeu in Australia, the apteryx, dinornis, etc., in New Zealand, the epiornis in Madagascar. Still the existence of such a land would not alone explain the various geographical cross-relations which have been given above. It would not, for example, account for the resemblance between the crustacea or fishes of New Zealand and of England. It would, however, go far to explain the identity (specific or generic) between fresh-water and other forms now simultaneously existing in Australia and South America, or in either or both of these, and New Zealand. Again, mutations of elevation small and gradual (but frequent and intermitting), through enormous periods of time—waves, as it were, of land rolling many times in many directions—might be made to explain many difficulties as to geographical distribution, and any cases that remained would probably be capable of explanation, as being isolated but allied animal forms, now separated indeed, but being merely remnants of extensive groups which, at an earlier period, were spread over the surface of the earth. Thus none of the facts here given are any serious difficulty to the doctrine of “evolution,” but it is contended in this book that if other considerations render it improbable that the manifestation of the successive forms of life has been brought about by minute, indefinite, and fortuitous variations, then these facts as to geographical distribution intensify that improbability, and are so far worthy of attention. All geographical difficulties of the kind would be evaded if we could concede the probability of the independent origin, in different localities, of the same organic forms in animals high in the scale of nature. Similar causes must produce similar results, and new reasons have been lately adduced for believing, as regards the lowest organisms, that the same forms can arise and manifest themselves independently. The difficulty as to higher animals is, however, much greater, as (on the theory of evolution) one acting force must always be the ancestral history in each case, and this force must always tend to go on acting in the same groove and direction in the future as it has in the past. So that it is difficult to conceive that individuals, the ancestral history of which is very different, can be acted upon by all influences, external and internal, in such diverse ways and proportions that the results (unequals being added to unequals) shall be equal and similar. Still, though highly improbable, this cannot be said to be impossible; and if there is an innate law of any kind helping to determine specific evolution, this may more or less, or entirely, neutralize or even reverse the effect of ancestral habit. Thus, it is quite conceivable that a pleurodont lizard might have arisen in Madagascar in perfect independence of the similarly-formed American lacertilia : just as certain teeth of carnivorous and insectivorous marsupial animals have been seen most closely to resemble those of carnivorous and insectivorous placental beasts; just as, again, the paddles of the Cetacea resemble in the fact of a multiplication in the number of the phalanges, the many-jointed feet of extinct marine reptiles, and as the beak of the cuttle-fish or of the tadpole resembles that of birds. We have already seen (in Chapter III.) that it is impossible, upon any hypothesis, to escape admitting the independent origins of closely-similar forms. It may be that they are both more frequent and more important than is generally thought. That closely-similar structures may arise without a genetic relationship has been lately well urged by Mr. Ray Lankester.” He has brought this notion forward even as regards the bones of the skull in Osseous fishes and in mammals. He has done so on the ground that the probable common ancestor of mammals and of Osseous fishes was a vertebrate animal of so low a type that it could not be supposed to have possessed a skull differentiated into distinct bony elements—even if it was bony at all. If this was so, then the cranial bones must have had an independent origin in each class, and in this case we have the most strikingly harmonious and parallel results from independent actions. For the bones of the skull in an osseous fish are so closely conformed to those of a mammal, that “both types of skull exhibit many bones in common,” though “in each type some of these bones acquire special arrangements and very different magnitudes.”” And no investigator of homologies doubts that a considerable number of the bones which form the skull of any osseous fish are distinctly homologous with the cranial bones of man. The occipital, the parietal, and frontal, the bones which surround the internal ear, the vomer, the premaxilla, and the quadrate bones, may be given as examples. Now if such close relations of homology can be brought about independently of any but the most remote genetic affinity, it would be rash to affirm dogmatically that there is any impossibility in the independent origin of such forms as centetes and solenodon, or of genetically distinct batrachians, as similar to each other as are some of the frogs of South America and of Europe. At the same time such phenomena must at present be considered as very improbable, from the action of ancestral habit, as before stated. We have seen, then, that the geographical distribution of animals presents difficulties, though not insuperable ones, for the Darwinian hypothesis. If, however, other reasons against it appear of any weight—if, especially, there is reason to believe that geological time has not been sufficient for it, then it will be well to bear in mind the facts here enumerated. These facts, however, are not opposed to the doctrine of evolution ; and if it could be established that closely-similar forms had really arisen in complete independence one of the other, they would rather tend to strengthen and to support that theory.

* See Ann, and Mag. of Nat. Hist., July, 1870, p. 37. * Prof. Huxley's Lectures on the Elements of Comp. Anat., p. 184.

CHAPTER VIII.

HOMOLOGIES.

Animals made up of Parts mutually related in Various Ways.--What Homology is. —Its Warious Kinds,--Serial Homology.--Lateral Homology.—Wertical Homology. —Mr. Herbert Spencer's Explanations.—An Internal Power necessary, as shown by Facts of Comparative Anatomy.—Of Teratology.—M. St. Hilaire.—Prof. Burt Wilder—Foot-wings.-Facts of Pathology.—Mr. James Paget.—Dr. William Budd.—The Existence of such an Internal Power of Individual Development diminishes the Improbability of an Analogous Law of Specific Origination.

THAT concrete whole which is spoken of as “an individual” (such, e. g., as a bird or a lobster) is formed of a more or less complex aggregation of parts which are actually (from whatever cause or causes) grouped together in a harmonious interdependency, and which have a multitude of complex relations among themselves.

The mind detects a certain number of these relations as it contemplates the various component parts of an individual in one or other direction—as it follows up different lines of thought. These perceived relations, though subjective, as relations, have nevertheless an objective foundation as real parts, or conditions of parts, of real wholes; they are, therefore, true relations—such, e. g., as those between the right and left hand, between the hand and the foot, etc.

The component parts of each concrete whole have also a relation of resemblance to the parts of other concrete wholes, whether of the same or of different kinds, as the resemblance between the hands of two men, or that between the hand of a man and the fore-paw of a cat.

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